John, an ambitious but undisciplined New York City office worker, meets and marries Mary. They start a family, struggle to cope with marital stress, financial setbacks, and tragedy, all while lost amid the anonymous, pitiless throngs of the big city.
Tillie is a secretary always dressed in the height of fashion who tries to capture a millionaire named Pennington Fish. Once she gets a stenographic position at Mr. Simpkins's company she sets her cap for the general manager, Benjamin Franklin Whipple. Eventually Tillie announces that she is going to "catch the rich Mr. Fish by using Whipple as the worm."
May s married to Roger, an alcoholic hell-raiser. During one of their riotous parties, she tests his fidelity by impersonating a notorious masked dancer and trying to seduce him.
Katherine Emerson, an Iowa girl hungry for the good things in life, leaves her small hometown and sets out for New York. En route, she is involved in a train wreck in which another woman is killed. Katherine finds the woman's purse and, among its contents, discovers an invitation for the woman to spend 6 months in an unoccupied luxury apartment in Manhattan. Katherine seizes this opportunity and sets up housekeeping in the elegant suite, living well and dressing in the newest fashions.
When Dorothy wants to marry Bob (Robert Agnew), her mother, Mildred, forbids the match. Dorothy angrily asserts that Mildred might reconsider if her own mother had forbid her marriage. The rest of the film is a flashback, as Mildred recalls her own youth, when her dictatorial mother did forbid her to marry Lyman. Lyman enlisted with Teddy Roosevelt's Rough Riders to fight in the Spanish-American War, but was killed in battle.
