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The Ark Fleet has been destroyed, and a significant number of the enemy's troops have been wiped out due to its crash landing. As the remaining forces of the Holy Order of Michael regroup in order to launch a final assault on Weiswolf Castle, the wZERO unit, along with their new ally Ashley Ashra, stand ready to intercept them. Meanwhile, with his Geass out of control, Shin moves to erase his younger brother's existence once and for all. But Akito, having promised Leila that he will come back alive, refuses to accept such a fate, and the two clash in their final battle.
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While W-0 is away, Leila and the remaining crew at base find themselves surrounded by Britannian Knightmares. Piloting the Vercingetorix, Shin Hyuuga leads the enemy charge, having usurped the position of Grand Master in the Knights of St. Michael. With her core members unavailable, it's up to Leila to protect her comrades, but this may be much tougher than she realizes: a spy lurking among them is finally ready to reveal himself. Left with no choice but to surrender, Leila arranges a meeting with Shin in order to ensure the safety of her squad. But as negotiations begin to fall apart, Akito and an unexpected ally return in the nick of time to turn things around.
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Os acontecimentos deste filme precedem os eventos da série, mostrando os estranhos acontecimentos por detrás da parceria entre Shinjuurou e Inga.
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Século Universal 0087, Char Aznable agora se tornou o líder do AEUG. Scirocco, o homem de Júpiter, planeja tomar o controle dos Titãs. Haman, o líder do Eixo, está tentando restaurar a família Zabi. Através da política e da estratégia, esses três poderes lutam pelo domínio sobre a esfera da Terra. A guerra se transforma em um final mortal em torno da colônia espacial Gryps 2, que foi convertida em um canhão laser gigante. Que destino espera Kamille no final do conflito? Esta é a parte final da trilogia de recursos do Mobile Suit Zeta Gundam, que apresenta animação aprimorada e músicas tema por GACKT.
