Yapool has returned after being injured in Ultraman Ginga S. He masterminds several attacks on Shou's homeworld, the Victorian kingdom in an attempt to get to the Victorium Core. Ultraman Hikari appears and grants Shou the ability to transform into Knight Mode with the Knight Timbre. Now free from their Spark Doll forms, some of the Showa Ultras join Victory in his fight.
Toron, a young tyrannosaurus boy, is carried off downstream and loses his memory. While making his way back home, he discovers new friends and finds out that working together is the only way forward in life.
One day, Mana Aida receives a wedding dress from her mother and grandmother that they had worn in the past. While thinking about who she would wear this beside someday, a mysterious man named Marsh appears, and Mana is thrown back into the past. She was happy to see her grandmother she loved and her old dog Maro, but she then realizes that Alice and Rikka aren't there because memories have been altered. In order to save precious memories, the future, dreams and hope, Mana transforms into Cure Heart.
