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2199 AD. Yamato tried to leave behind the Large Magellanic Cloud galaxy, after receiving a "Cosmo Reverse System" at its destination, Iscandar. However, suddenly, it encounters a mysterious group at the edge of the Large Magellanic Cloud. The crew finds out that the group is called the "Gatlantis," and that its leader is the Gutaba expeditionary force commander who calls himself "Goran Dagaamu of Thunder." The commander demands that Yamato be handed over to him. Yamato, wanting to hurry to Earth, escapes the fray. However, Yamato is attacked by a devastating beam weapon called the "Flame Direct Attack Cannon" that can transcend space.
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В центре сюжета - начало карьеры офицера Альянса Джеймса Веги. Джеймс приходит в элитный отряд спецназа Альянса в начале борьбы с таинственной угрозой, известной как Коллекционеры. Состоя на службе в отдалённой звёздной системе, Джеймс Вега и его напарники должны защитить граждан колонии от вторжения неизвестных, которые пытаются похитить людей.
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Zephyr, now known as Z, rides the seas with only one goal: Destroy all pirates and their dreams at becoming King of Pirates. When Luffy and his crew encounter him at sea, not only are they utterly defeated by the man with an arm made of Seastone, Nami, Robin, and Chopper are turned 10 years younger due to Z's minion Ain. Luffy is so determined to win against him that he does not even notice Z's master plan that could sacrifice thousands of lives.
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Фильм повествует о пирате, бывшим главным противником Гола Д. Роджера — Шики «Золотым Льве». Он крадёт у Мугивар Нами и просит её вступить в его команду. Остров с загадочными животными. Коварные планы Шики. Что ждёт Мугивар в поисках Нами?
