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ゴジラシリーズ第26作。「ミレニアムシリーズ」第4作。1954年、ゴジラ日本上陸。絶体絶命の中、芹沢博士が開発した特殊兵器オキシジェン・デストロイヤーでゴジラを撃退。だが、博士はこの兵器に脅威を感じ、自らの命と共に消滅させる。この年、政府は対特殊生物防衛専門の“特生自衛隊”を組織。99年、再びゴジラ出現。特生自衛隊は迎撃するも苦戦を強いられる。政府は人工生物学者・湯原徳光を中心とする科学者チームへ対G兵器・機龍“メカゴジラ”開発を要請。2003年、ついにメカゴジラが完成する。そして、機龍の操縦士として、99年の戦いで仲間を失った特生自衛隊員・家城茜が任命された…。
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The hero Nike and magic user Kukuri learn of the "Pickle of Happiness", an item that will grant a wish to the one who eats it. With the help of old friends, they must beat the monsters to the Pickle and convince its guardian dragon to give it to them.
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In the future, the oceans have risen to flood all the continents due to humanity's negligence of the environment. Only a few bits of land have been spared, and one of those is the city-island where Elfie lives with her grandfather. One day, due to an underwater incident, she discovers that she can breathe underwater. Her grandfather reveals that in truth she is one of the mythical sea-people. Using the city folk's xenophobia, local politicians spark a war with this hidden people to distract people from their current resource problems, and Elfie is caught in the middle of it.
